LOS ANGELES -- After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

San Diego’s “Queen of the Zoo” has died. Her name was Gramma, and she was a Galápagos tortoise whose 141 years of existence (give or take) included more than 20 presidents, two world wars and a pair ...

(KSWB/KUSI) — Gramma, a Galapagos tortoise that was the oldest living animal at the San Diego Zoo, has died. The zoo said Gramma had ongoing bone conditions and that caretakers had to make the ...